Exploring Georgia's Natural History
  • Home: Georgias Fossils
  • 1: Georgia's Oldest Fossils; Archaeocyathids, At 513 Million Years Old
  • 2: Trilobites; 500 Million Years Ago
    • 2A; Murray County Stromatolites
    • 2B; A Trilobite Nest in Georgia
  • 3: Geologic Time
  • 4: Georgia's Oldest Vertebrate?
  • 5: NW Georgia, 488 to 300 million years ago
    • 5A; Georgia’s Pennsylvanian Plant Fossils
    • 5B: Carpentertypus durhami, Georgia’s Giant Insect, 315 Million Years Ago
    • 5C: Mississippian Trilobites in Northwest Georgia Describing the New Species Australosutura georgiana
    • 5D: Crinoids & Blastoids Of Northwest Georgia
    • 5E; Fossil Locations of Northwest Georgia
    • 5F: Pennsylvanian Plant Fossils of NW Georgia
    • 5G; Ordovician Invertebrates of Northwest Georgia
    • 5H: Trace Fossils in NW Georgia’s Metamorphic Rock
  • *NEW* 5I; Georgia’s Oldest Shark Tooth
  • 6: 200 Million Years Ago
    • 6A: Birth of the Atlantic Ocean
  • 7: Cretaceous Georgia, Dinosaurs & more
    • 7A: Georgia's Pterosaur
    • 7B: So Many Sharks
    • 7C: Coelecanths
    • 7D: Xiphactinus vetus
    • 7E: Side-necked turtles
    • 7F: Marine Reptiles
    • 7G: Dinosaurs in Georgia
    • 7H: Deinosuchus schwimmeri in Recognition of Dr. David Schwimmer
    • 7I; The Blufftown Formation
    • 7J: New Species of Cretaceous Flowers Reported From Crawford County
    • 7K: Field Trip, Chattahoochee River Valley 1980
    • 7L: The Eutaw Formation
    • 7M: The Pio Nono Formation
    • 7N: Plant Fossils of Crawford County, GA
    • 7O; 1914 Report Georgia Plant Fossils From the Upper Cretaceous
    • 7P: Bill Montante's Mega "Gator" Tooth Discovery
  • 8: Suwannee Current, Gulf Trough, & Bridgeboro Limestone
  • 9: 60 million years ago, The Paleocene's Clayton Formation, A Report; By Hank Josey
    • 9A: The Georgia Turtle
    • 9B; Sassafras Hill Quarry Huber Formation Plant Fossils in Kaolin
  • 10: The Eocene; Georgia's Oldest Mammals
    • 10A: The Origins of Whales
  • 11: A Whale For Georgia
  • 12: Basilosaurids; The First Modern Whales
    • 12A: Basilosaurus cetoides
    • 12B: Basilotritus
    • 12C: Crisp County & Cynthiacetus
    • 12D: Chrysocetus
    • 12E: The Redmond Mandible of Albany Ga
    • 12F: The Tivola Whale; From Houston County to the Smithsonian
  • 13: Ziggy and The Museum of Arts & Sciences, Macon, GA
  • 14: Late Eocene
    • 14A: Eocene Fossils & Stratigraphy
    • 14B; Fossils, Impacts, & Tektites Dating the Clinchfield Formation
    • 14C: The Tivola Limestone
    • 14C1: Oldest Oreodont in the Southeast & Georgia's first!
    • 14D: Twiggs Clay Vertebrates
    • 14E: Ocmulgee Formation Vertebrates
    • 14F; Sandersville Limestone, By Hank Josey
    • 14I: Dating Late Eocene Sediments
    • 14J: Georgia's Tektites; Georgiaites
    • 14K; Shell Bluff; Georgia's Most Historic Paleontology Site
    • 14L; Taylors Bluff, Paleo Paddling the Ocmulgee River
    • 14M; Eocene Terrestrial Mammals From Gordon, GA
    • 14N: Fossil Ridge, A Stratigraphic Study in Oaky Woods Wildlife Management Area
    • 14O; Georgia's First Entelodont
    • 14P: Historic Rich Hill
    • 14Q; Bibb County's Christy Hill, Clinchfield Formation Hilltop
    • 14R: Browns Mount, The Fall Line, Elevations, Uplifts, & Native Middle Georgians
  • 15: Early Oligocene
    • 15A: The Marianna Limestone
    • 15B; The Glendon Limestone
    • 15C: Undiffereniated Oligocene Residuum
    • 15D; Brissus bridgeboroensis; A New Echinoid Species From Georgia’s Bridgeboro Limestone
    • 15E: The Curious Steinkern Sea Biscuits of Red Dog Farm Road
    • 15F: Early Oligocene Gordian Knot
  • 16: Bonaire GA Entelodont
  • 17: The Whale Eating Shark
  • 18: Miocene Epoch; 23.3 to 5.3 Million Years Ago
    • 18A; Miocene Terrestrial Vertebrates
    • 18B; Paul Fell, Rockhouse Cave
    • 18C: The Marks Head Formation
    • 18D: Miocene Terrestrial Vertebrates of the Marks Head Formation
    • 18E: The Statenville Formation
    • 18F: Georgia’s 13 Million Year Old Dugong Metaxytherium calvertense
    • 18G: Gastropod Gulch, Julia Gardner, & Miocene Invertebrates In Decatur County
    • 18H; Bony Bluff, Rocky Ford, Echols County In Southernmost Georgia
    • 18I: Decatur County Fossils & Natural History
    • 18J; Decatur County Vertebrates, FLMNH
    • 18K; A Large Tegu-Like Lizard, Middle Miocene, Climate Optimum, Decatur County, GA
  • 19: Pliocene Epoch; 5.3 to 2.5 Million years Ago
    • 19A: Two Small Primitive Horses from Taylor County Advance the Science of Georgia Geology
  • 20: The Pleistocene & Holocene Epochs, The Ice Ages
    • 20A; Clark Quarry's Mammoths & Bison
    • 20B: Pleistocene Vertebrate List
    • 20C: Georgia’s Eolian Dunes
    • 20D: Georgia’s Carolina Bays
    • 20E: Late Pleistocene Significant Events
    • 20F: Southeastern Thermal Enclave
    • 20G; Diamond Back Terrapins
    • 20H; A Kaolin Mine Beaver Dam
    • 20I; Pleistocene Vertebrate Fossils On Georgia’s Piedmont
    • 20J; Watkins Quarry Pleistocene Vertebrates, Glynn County, GA
    • 20K: Pleistocene Vertebrates from Coastal Georgia
    • 20L; Sandy Run Creek Core, Warner Robins, Houston County, GA
    • 20M: Bone Bed, Pleistocene, Coastal Georgia
    • 20N: Caribou & Elk Fossils from Georgia & Alabama
    • 20O; Tapir Veroensis, Walker County, Late Pleistocene
    • 20P; Ladds , School Teachers, Pleistocene Vertebrates, Bartow County, GA
    • 20Q; Chet Kirby, Glynn County, GA Pleistocene & Miocene-Pleistocene
  • 21: Humans in Georgia
  • 22A: Echinoids of Georgia, Cenozoic Era (Sand Dollars & Urchins)
    • 22B: Echinoids of Georgia, Cenozoic, By County
  • 23A; Exploring the Paleontology of Southernmost Georgia
    • 23B; Seminole County
    • 23c: Grady County Blowing Caves, Forest Falls, Fossils & Natural History
  • 24: Georgia's Meteorites
    • 24A: Did I Find A Meteorite?
    • 24B: Georgia's Lost Meteorite
    • 24C: The Sardis Iron, Georgia's Largest Meteorite
  • 25: Dr. Burt Carter, Georgia Southwestern, Professor Invertebrate Paleontologist, Emeritus
    • 25A; Burt Carter, Uniformitarianism
    • 25B; Burt Carter, Inclusions
    • 25C; Burt Carter, Superposition
    • 25D; Burt Carter, Principal of Horizonality
    • 25E, Burt Carter, Cross Cutting
    • 25F; Burt Carter, Deep Time
    • 25G; Burt Carter, Fossil Succession
  • 26: Paul F. Huddlestun Coastal Plain Core Logs
    • 26A: Late Eocene & Older... Coastal Plain Stratigraphy
    • 26B: Gulf Trough Cores, Colquitt County, by Paul Huddlestun
    • 26C; Washington County Core Logs By Paul Huddlestun
    • 26D: Coastal Plain Core Logs by Paul F. Huddlestun
  • 27: Science, Georgia Research
    • 27A: Coastal Plain Correlation Chart
    • 27B: Physiographic Map of Georgia
    • 27C: Collections & Stewardship of Georgia’s Fossils
    • 27D: Needed; The Georgia Geologic Survey
    • 27E: GA County Localities, Houston County
    • 27F: Trace Fossils on the Coastal Plain
    • 27G: Georgia’s Decapod Fossils
    • 27H: Georgia Vertebrates in the Florida Museum of Natural History
  • 28: Educational Matetrial For Georgia Classrooms
    • 28A: Oaky Woods Stratigraphy, PowerPoint
    • 28B: Fossils of Oaky Woods
    • 28C: I, Periarchus (A Fossil's Tale)
    • 28D: The Tivola Whales (April 2023 talk to the Mid-Georgia Gem & Mineral Society)
    • 28E: Georgiacetus Presentation; A Whale for Georgia
    • 28F: My Field Kit; What You Need In The Field
    • 28G: Meet Crassostrea gigantissima, Georgia's Historic Giant Oyster
    • 28H: The Natural History & Fossils Record of Houston County, GA
    • 28I: Evidence for Evolution in Georgia's Fossil Record... A look at Teeth
    • 28J: Georgia's State Fossil; Shark Teeth
    • 28K; An Introduction To Fossils; Presentation
    • 28L: Library & School Presentations
    • 28M: Georgia's Paleontology For Georgia's Classrooms

18K; A Large Tegu-Like Lizard.
Middle Miocene, Climate Optimum,
Decatur County, GA

By
Thomas Thurman
02/March/2026
 

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​A large Tegu-like lizard hunted Decatur County in Southwest Georgia about 14.4 million years ago. In 2025 Jason R. Bourque and Edward L. Stanley announced a new genus and species of lizard known only from Decatur County, Georgia, Wautaugategu formidus. The fossil was a single thoracic vertebra. Their research appeared in the prestigious Journal of Paleontology. Their paper can be found at the base of this page. 
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Decatur County about 14.4 million years ago was a very different place. Sea levels were high and Florida was generally submerged with only an island chain, the Brooksville Ridge, above sea level. Decatur County was coastal and on the edge of the paleo Flint River, Chattahoochee River, and Spring Creek deltas. It isn’t impossible that the Paleo-Flint River met the sea very near where Gragg Mine stood.
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Today, the confluence of these waterways is dammed at the Florida line by the Woodruff Dam. But if you look today at Google Earth and check the elevations around Gragg Mine you’ll see that it’s on modestly higher ground today that most of the surrounding area. Was this true during the Miocene? That’s hard to say, but it does suggest that this area could have been an island during high stands of sea levels.  
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​Strange Gomphotherium elephants walked the county, they had two short tusks pointing down from the lower jaw and two larger tusks in the upper jaw, also pointing down with a rather short elephant trunk. 
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​The large carnivore Amphicyon ingens or bear-dog was also present in numbers. A powerful, robust hunter whose shoulders were 1.25 meters (4+ feet) tall and whose dog-like head was as big as a man’s chest. 
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​There were rhinoceroses, populations of early horses, populations of deer-like (Genus Paratoceras & family Dromomerycidae) animals whose antlers were fur covered, bony extensions of their skulls. Giant tortoises (Hesperotestudo), peccaries, wild dogs, members of the camel family, beavers, rabbits and many others. Decatur County was a very different place. Fossils from all of these were found within a few miles of Attapulgus. 
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Lists of recovered Miocene fossils can be found on this website on page #18J; Decatur County, Georgia’s Miocene Vertebrates In The Florida Museum of Natural History.
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I want to thank Rachel Narducci, Lead Collections Manager of Vertebrate Paleontology at the Florida Museum of Natural History for forwarding this paper to me.
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The Lizard
Wautaugategu formidus, is a member of the Tupinambinae (Teiidae family). Bourque and Stanley tell us that the Tupinambinae are group of medium to large bodied lizards native to South America which includes Tegu lizards which grow up to 1.3 meters (4 feet) long. The Tupinambinae are well represented in South America from the Eocene to the Pleistocene and believed to be more widespread in the Mesozoic to Paleogene. But this vertebra is the “oldest unequivocal record” in North America.
  
Etymology
To quote the authors: Combination of Wautauga in reference to the Wautauga State Forest, a local landmark near the type locality, and tegu, the colloquial name for T. teguixin and Salvator spp.”

“Wautauga State Forest” only comes up in a google search under the Climax, Georgia Chamber of Commerce. I see no listing for it in the Georgia Department of Natural Resources or U.S. Forest Service. Nor does it come up in Google Earth, though Google Earth produces two references on the name, one in West Virginia and one in Tennessee. Google Earth also shows a Wautauga Road running NE out of Attapulgus, Georgia but it’s over six miles from Climax at its closest point.  
 
The fossil vertebra establishing Wautaugategu formidus was produced by Gragg Mine which is 3.5 miles SE of Attapulgus and more than 3 miles south of Wautauga Rd. So I’m not sure about the etimology of Wautauga
 
The specific name, formidus, is a Latin word meaning 'warm', alluding to the climatic conditions the taxon would have inhabited and the preferred body temperatures of living relatives.


Geologic Setting
Gragg Mine
The Wautaugategu formidus vertebra is Catalog# 546657 in the FLMNH database which can be accessed here; Web Portal. It came from the Gragg Mine, a now defunct fuller’s earth, clay mine. The fossil was collected in May 2005 by Jonathan Hoffman and Andrew Zimmerman. The Gragg Mine was located about 5.6 km southeast of Attapulgus but has since been reclaimed and reported from 2023 say the fossil beds are reforested, inaccessible, and likely buried during reclamation of the mine.
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The FLMNH database states the fossil came from the Torreya Formation, which is middle Miocene, but the paper does not state that.
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​Bourque and Stanley dated the sediments biochronologically, meaning by the presence of identified fossils whose range in geologic time is well known and established. Many fossils attributed to this, many mammals were recovered and identified and their fossils helped constrain the age of the deposit. Other fossils also played their part, particularly the presence of the rare, large-bodied salamander Batrachosauroides dissimulan (catalog# 217589) which exceeded three feet helped define the age of the sediments. Multiple specimens of this salamander genus were found in Gragg Mine and the Florida Museum holds other specimens of the same species from Florida and Texas, so the salamander could be confidently identified. Collectively the fossils established the age of the sediments as Middle Miocene, NALMA Barstovian 2, 14.8 to 14.0 million years old. (NALMA; North American Land Mammal Age)
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Environment
The sediments represent a coastal fluvial deposit (Fluvial, river or stream sourced) with a mixture of marine, freshwater, and terrestrial vertebrates. That’s the paleo-environment. Gragg Mine is just south of the Flint River.
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​​These fossils are likely the remains of animals which got caught in ancient rivers, most likely the Paleo Flint River  The Gragg Mine fossils included shark teeth, turtle material, snake material, lizards, Amphiuma, frogs, & others. “Marine and freshwater fish are the most abundant non-mammalian constituents.” (Bourque & Stanley) Another small-bodied teiid lizard was also present. 
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The Gragg Mine site is regionally important as it represents a coastal plain at the conclusion of one of the warmest periods of the Middle Miocene Climatic Optimum when sea levels were at their highest for the Neogene Period; 2.58 to 23.03 million years ago. As mentioned earlier, Florida was submerged except for the Brooksville Ridge in northern peninsular Florida as a series of islands and a scattering of barrier islands along the modern Georgia, Alabama border with Florida.
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Bourque & Stanley's Paper
bourque_stanley_tegulike_lizard_gragg_quarry.pdf
File Size: 9911 kb
File Type: pdf
Download File

Reference:
  • Bourque, Jason R. & Stanley, Edward L.; A Tegu-Like (Teiidae, Tupinambinae) from the Middle Miocene Climatic Optimum of the Southeastern United States. Journal of Paleontology, 99(1), pages 177-191, 2025